烟管蜗牛科
Clash Royale CLAN TAG#URR8PPP | 烟管蜗牛科 | |
|---|---|
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Cochlodina laminata | |
科学分类  | |
| 界： | 动物界 Animalia |
| 门： | 軟體動物門 Mollusca |
| 纲： | 腹足纲 Gastropoda |
| 目： | 柄眼目 Stylommatophora |
| 亚目： | 旋蝸牛亞目 Helicina |
| 下目： | 烟管蜗牛下目 Clausilioidei |
| 总科： | 烟管蜗牛总科 Clausilioidea |
| 科： | 烟管蜗牛科 Clausiliidae J. E. Gray, 1855[1] |
模式属 | |
烟管蜗牛属 Clausilia | |
烟管蜗牛科（學名：Clausiliidae），亦作烟管螺科[2]或密封蝸科，是烟管蜗牛总科之下的一種小型和非常長的科級分類。本科物種主要都是左旋物種的陸生蝸牛、會呼吸空氣的有肺類腹足綱直神經亞綱柄眼目軟體動物[3]:105。
本科舊屬柄眼目中尿道亞目（Mesurethra）[4]及泛有肺類真有肺類柄眼類非正式群組弯尿道類。
截至2008年10月 (2008-10)[update]，本科連同現生種及化石種合共有1300個物種，是陸生蝸牛當中其生物多樣性最高的分類單元（參照Orthalicidae，在整個腹足綱來說也只是僅次於海生的小塔螺科）。
本科物種在台灣的名稱叫作「烟管蝸牛科」，這是得名於其殻口的獨特「烟管」解剖結構，叫作「腔襞」[5]:116，讓牠們可以將牠們的口蓋變得可以像一扇趟門那樣把殻口關閉起來。
目录
1 型態描述
1.1 「烟管」
2 生物學
3 分類學
3.1 2005年分類
3.2 2017年分類
4 屬
4.1 烟管蜗牛亚科
4.2 Alopiinae
4.3 Baleinae
4.4 Constrictinae
4.5 斜管螺亞科
4.6 Eualopiinae
4.7 Laminiferinae
4.8 Mentissoideinae
4.9 大口管螺亚科
4.10 管螺亚科
4.11 Serrulininae
5 保育
6 參考文獻
7 延伸閱讀
8 外部連結
型態描述
Megalophaedusa martensi is the largest species of the family Clausiliidae in the world. The individual in the photograph has a shell which is 45.5 mm long. The whole snail weighs 3.2 g live.
幾乎所有本科的貝殻都是左旋的[3]:105，這在腹足綱物種的螺殻來說並不常見。殻的表面有肋樣構造[3]:105。殻長約10到15 mm[3]:105。
These snails have shells which are extremely high-spired, with numerous whorls.
The shells tend to be club-shaped, tapering at both ends to a rounded nub. The aperture usually has visible folds.
「烟管」
Clausilia dubia的「烟管」構造。
烟管蜗牛科物種在肺螺類腹足綱軟體動物來說相當特殊，因為牠們大多數的殻口都有稱為「腔襞」的類似「烟管」構造，就像一扇趟門一樣。這種「烟管」與口蓋並不一樣，因為在肺螺類物種都已有外套膜肺，並沒有「口蓋」這種構造。
腔襞是一種石灰質構造，舌形或調匙形，可將其殻口完全關閉，以防止肉食性的甲蟲幼蟲入侵其柔軟的身體。腔襞有不同的形態，如「月形」[5]:116或「弓形」[6]腔襞、「倒C形腔襞」[5]:124、「工形腔襞」[5]:118、「平行腔襞」[5]:128等，還可以分為多個部分，如：上腔襞、下腔襞、月形襞及主襞等[5]:116。腔襞有「閉板」及「腔襞板」（clausilium slides）將其封閉：「閉板」是附生於殻軸的板狀構造，而腔襞板則是殻內壁的板狀突起[5]:116，由殻壁摺疊而形成。
生物學
本科物種的染色體的單倍體介乎21到30之間[7]:139,142。
分類學
本科的模式屬是烟管蜗牛属（Clausilia Draparnaud, 1805）。
根據2005年《布歇特和洛克羅伊的腹足類分類》，本科是彎尿道類非正式群組的成員[8]，而這個群組在2010年的修訂被歸類為泛有肺類真有肺類演化支的成員[9]。在2017年《布歇特等人的腹足類分類》，曲尿道類支序和泛有肺類都不再使用[10]。
2005年分類
根據2005年《布歇特和洛克羅伊的腹足類分類》，本科包括下列各亞科：
烟管蜗牛亚科 Clausiliinae Gray, 1855- tribe Clausiliini Gray, 1855 - synonym: Fusulinae Lindholm, 1924
- tribe Gracillariini H. Nordsieck, 1979
- subfamily Alopiinae A. J. Wagner, 1913
- tribe Alopiini A. J. Wagner, 1913
- tribe Cochlodinini Lindholm, 1925 (1923) - synonym: Marpessinae Wenz, 1923
- tribe Delimini Brandt, 1956 - synonym: Papilliferini Brandt, 1961 (n.a.)
- tribe Medorini H. Nordsieck, 1997
- tribe Montenegrinini H. Nordsieck, 1972
- subfamily Baleinae A. J. Wagner, 1913 - synonyms: Laciniariini H. Nordsieck, 1963; Tristaniinae Schileyko, 1999
- subfamily † Constrictinae H. Nordsieck, 1981[11]
斜管螺亞科 Garnieriinae C.R. Boettger, 1926
斜管螺族 Garnieriini C.R. Boettger, 1926
伞管螺族 Tropidaucheniini H. Nordsieck, 2002
- subfamily † Eualopiinae H. Nordsieck, 1978
- tribe † Eualopiini H. Nordsieck, 1978[12]
- tribe † Rillyini H. Nordsieck, 1985[13]
- subfamily Laminiferinae Wenz, 1923
- subfamily Mentissoideinae Lindholm, 1924
- tribe Mentissoideini Lindholm, 1924 - synonym: Euxininae I. M. Likharev, 1962
- tribe Acrotomini H. Nordsieck, 1979
- tribe Boettgeriini H. Nordsieck, 1979
- tribe Euxinellini Neubert, 2002
- tribe Filosini H. Nordsieck, 1979
- tribe Olympicolini Neubert, 2002
- tribe Strigileuxinini H. Nordsieck, 1994
- tribe Strumosini H. Nordsieck, 1994
- subfamily Neniinae Wenz, 1923 - Neniastrinae H. B. Baker, 1930
- subfamily Phaedusinae A. J. Wagner, 1922
- tribe Phaedusini A. J. Wagner, 1922
- tribe Megalophaedusini Zilch, 1954 - synonym: Zaptyxini Zilch, 1954
- subfamily Serrulininae Ehrmann, 1927
2017年分類
根據2017年《布歇特等人的腹足類分類》，煙管蝸牛科是柄眼目旋蝸牛亞目煙管蝸牛下目之下的煙管蝸牛總科的成員。本科之下的亞科從過往的11個亞科縮減至8個亞科21個族，分別如下：
- Subfamily Alopiinae A.J. Wagner, 1913
- Tribe Alopiini A.J. Wagner, 1913
- Tribe Cochlodinini Lindholm, 1925 (1923)
- Tribe Delimini R. Brandt, 1956
- Tribe Medorini H. Nordsieck, 1997
- Tribe Alopiini A.J. Wagner, 1913
- Subfamily Clausiliinae Gray, 1855
斜管螺亞科 Garnieriinae C.R. Boettger, 1926- Subfamily Laminiferinae Wenz, 1923
- Subfamily Mentissoideinae Lindholm, 1924
- Subfamily Neniinae Wenz, 1923
- Subfamily Peruiniinae H. Nordsieck, 2005
- Subfamily Phaedusinae A.J. Wagner, 1922
- Tribe Phaedusini A.J. Wagner, 1922
- Tribe Serrulinini Ehrmann, 1927
- Tribe Synprosphymini H. Nordsieck, 2007
- Tribe Phaedusini A.J. Wagner, 1922
屬
以下詳列本科之下各個亞科所屬的分類單元：
烟管蜗牛亚科
烟管蜗牛亚科（Clausiliinae）包括以下兩族：
烟管蜗牛族（Clausiliini）
烟管蜗牛属 Clausilia Draparnaud, 1805 - with two subgenera: Clausilia and Strobeliella H. Nordsieck, 1977[14]
Erjaveci a Brusina, 1870[14]
Fusulus Fitzinger, 1833 - with two subgenera: Erjaveciella H. Nordsieck, 1877 and Fusulus[14]
Julica H. Nordsieck, 1963[14]
Macrogastra Hartmann, 1841 - with three subgenera: Macrogastra, Pseudovestia H. Nordsieck, 1877 and Pyrostoma Vest, 1867[14]
Micridyla H. Nordsieck, 1973[14]
Neostyriaca A. J. Wagner, 1920[14]
Pseudofusulus H. Nordsieck, 1977 - has the only one species: Pseudofusulus varians[14]
Ruthenica Lindholm, 1924[14]
- tribe Gracillariini
Graciliaria Bielz, 1867[15]
Alopiinae
Alopiinae[16]
tribe Alopiini
Alopia H. & A.Adams, 1855 - with two subgenera Alopia and Kimakowiczia Szekeres, 1969
Herilla H. & A. Adams, 1855
Triloba Vest, 1867
tribe Cochlodinini
Cochlodina A. Férussac, 1821 - with four subgenera: Cochlodina; Cochlodinastra H. Nordsieck, 1977; Paracochlodina H. Nordsieck, 1969; Procochlodina H. Nordsieck, 1969
Macedonica O. Boettger, 1877
tribe Delimini
Barcania Brandt, 1956
Charpentieria Stabille, 1864 - with four subgenera: Charpentieria; Gibbularia Monterosato, 1908; Mauritanica O. Boettger, 1877; Siciliaria Vest, 1867 and Stigmatica O. Boettger, 1877
Delima Hartmann, 1842 - with four subgenera: Delima; Dugiana Stamol & Slapnik, 2002; Piceata O. Boettger, 1877 and Semirugata O. Boettger, 1877
Dilataria Vest, 1867
Papillifera Hartmann, 1842
tribe Medorini H. Nordsieck, 1997
Agathylla H. & A.Adams, 1855 - with two subgenera Agathylla and Agathyllina H. Nordsieck, 1969
Albinaria Vest, 1867 - also include former genera Carinigera Moellendorff, 1873; Cristataria Vest, 1867; Isabellaria Vest, 1867; Sericata O. Boettger, 1878.
Inchoatia Gittenberger & Uit de Weerd, 2006[17]
Lampedusa - with two subgenera Imitatrix Westerlund, 1884 and Lampedusa
Leucostigma A. J. Wagner, 1919
Medora H. & A. Adams, 1855
Muticaria Lindholm, 1925
Strigilodelima A. J. Wagner, 1924
tribe Montenegrinini
Montenegrina O. Boettger, 1877
Protoherilla A. J. Wagner, 1921
Baleinae
Alinda H. & A. Adams, 1855 - with two subgenera: Alinda and Pseudalinda O. Boettger, 1877
Balea J. E. Gray, 1824
Bulgarica O. Boettger, 1877 - with three subgenera Bulgarica, Pavlovicia H. Nordsieck, 1973 and Strigilecula Kennard & Woodward, 1923
Lacinaria Hartmann, 1842
Likharevia H. Nordsieck, 1975
Menissa H. & A.Adams, 1855
Mentissella H. Nordsieck, 1973
Micropontica O. Boettger, 1881 - with two subgenera: Baleopsina Lindholm, 1924 and Micropontica
Mucronaria O. Boettger, 1877 - with two subgenera: Index O. Boettger, 1877 and Mucronaria
Quadriplicata O. Boettger, 1878
Vestia Hesse, 1916 - with three subgenera: Brabenecia H. Nordsieck, 1974; Vestia and Vestiella H. Nordsieck, 1877
Constrictinae
Fossil subfamily Constrictinae contains genera:
- † Constricta O. Boettger, 1877 - type genus of the subfamily[8]
斜管螺亞科
斜管螺亞科（Garnieriinae）包括下列各屬：
Garnieria Bourguignat, 1877
斜管螺属 Grandinenia H. Minato & C. Chen, 1984[18]
Indonenia Ehrmann, 1927
Megalauchenia H. Nordsieck, 2007
Neniauchenia H. Nordsieck, 2002
伞管螺属 Tropidauchenia Lindholm, 1924- subsp. Euryauchenia H. Nordsieck, 2007
- subsp. Tropidauchenia
Eualopiinae
Fossil subfamily Eualopiinae contains genera:
tribe † Eualopiini
- † Eualopia O. Boettger, 1877 - type genus of the subfamily[8]
tribe † Rillyini
- † Rillya Munier-Chalmas [in P. Fischer], 1883 - type genus of the tribe[8]
Laminiferinae
Laminifera O. Boettger, 1863 - the type genus of the family Laminiferinae.[8] Its type species is fossil.[8] (Actual status of the genus?)
Bofilliella Ehrmann, 1927
Neniatlanta Bouguignat, 1876
Mentissoideinae
Acrotoma O. Boettger, 1881 - with four subgenera: Acrotoma, Acrotomina H. Nordsieck, 1977; Bzybia H. Nordsieck, 1977 and Castelliana Suvorov, 2002
Akramowskia H. Nordsieck, 1975
Armenica O. Boettger, 1877 - with two subgenera: Armenica and Astrogena Szekeres, 1970
Bitorquata O. Boettger, 1877
Boettgeria O. Boettger, 1863 - with two subgenera: Boettgeriaand Loosjesiella Neubert & Groh, 1998
Elia H. & A.Adams, 1855 - with four subgenera: Acroeuxina O. Boettger, 1877; Caucasica O. Boettger, 1877; Elia and Megaleuxina O. Boettger, 1877
Euxina O. Boettger, 1877 - with two subgenera: Euxina and Illunellaria Lindholm, 1924
Euxinastra O. Boettger, 1888 - with two subgenera: Euxinastra and Odonteuxina H. Nordsieck, 1875
Euxinella H. Nordsieck, 1973
Filosa O. Boettger, 1877
Galeata O. Boettger, 1877
Idyla H. & A. Adams, 1855 - with two subgenera: Idyla and Strigilidyla H. Nordsieck, 1994
Kazancia Neubert, 1992 - with two subgenera: Kazancia and Lasica Neubert, 1995
Macroptychia O. Boettger, 1877
Mentissoidea O. Boettger, 1877
Olympicola Hesse, 1916
Phrygica H. Nordsieck, 1994
Roseniella Thiele, 1931 - with two subgenera: Chavchetia Neubert, 1992 and Roseniella
Sabaeola Lindholm, 1925
Sprattia O. Boettger, 1883
Strigileuxina H. Nordsieck, 1975
Strumosa O. Boettger, 1877
Sumelia H. Nordsieck, 1994
大口管螺亚科
大口管螺亚科（Neniinae Ehrmann, 1927）主要分佈於南美洲，但亦有少数分佈於中国南部、南亚及东南亚，包括以下各屬[19]：
Andinia Polinski, 1922
Andiniella Weyrauch, 1958
Audiniastra H. Nordsieck, 2005
Bequaertinenia Weyrauch, 1964 - with two subgenera: Bequaertinenia and Miranenia Grego & Szekeres, 2004
Brevinenia Neubert & H. Nordsieck, 2005
Columbinia Polinski, 1924 - with three subgenera: Columbinia; Paranenia Rehder, 1939 and Steatonenia Pilsbry, 1926
Cyclonenia H. Nordsieck, 1999
Cylindronenia Ehrmann in Pilsbry, 1949 - with two subgenera: Cylindronenia and Cylindroneniella H. Nordsieck, 2007
Ehrmanniella Zilch, 1949
Gonionenia Pilsbry, 1926
Gracilinenia Polinski, 1922
Hemicena Pilsbry, 1949
Incaglaia Pilsbry, 1949 - with two subgenera: Gibbonenia Zilch, 1954 and Incaglaia
Incania Polinski, 1922
Leuconenia H. Nordsieck, 2005
大口管螺屬 Nenia H. & A. Adams, 1855
Neniaptyx H. Nordsieck, 2007
Neniatracta Pilsbry, 1926
Neniella Grego & Sekeres, 2004
Neniops Pilsbry, 1926
Nenisca Rehder, 1939
Parabalea Ancey, 1882
Peruinia Polinski, 1922
Pfeifferiella Weyrauch, 1957
Pseudogracilinenia Loosjes & Loosjes-van Bemmel, 1984
Steeriana Jousseaume, 1900
Symptychiella H. Nordsieck, 1999 - with two subgenera: Divnenia H. Nordsieck, 2005 and Symptychiella
Temesa H. & A. Adams, 1855
Zilchiella Weyrauch, 1957
管螺亚科
管螺亚科（Phaedusinae Thiele, 1931）包括以下各屬[20]：
管螺族 Phaedusini
管螺属 Phaedusa H. & A.Adams, 1855 - with four subgenera: Metaphaedusa H. Nordsieck, 2001; Phaedusa; Pseudophaedusa Tomiyama, 1984 and Stereophaedusa O. Boettger, 1877
tribe Megalophaedusini
Leptacme Ehrmann, 1927[21]
Oospira Blanford, 1872[21] - with five subgenera: Atractophaedusa Ehrmann, 1927; Formosana O. Boettger, 1877; Formosanella H. Nordsieck, 2003; Oospira; Paraformosella H. Nordsieck, 2003 and Siphonophaedusa Lindholm, 1924
Zaptyx Pilsbry, 1901 - with two subgenera: Prozaptyx Loosjes, 1950 and Zaptyx
tribe ?
Acanthophaedusa H. Nordsieck, 2007
Bathyptychia Lindholm, 1925 - with three subgenera: Bathyptychia; Brachyptychia H. Nordsieck, 2001 and Strictiphaedusa H. Nordsieck, 2001
Celsiphaedusa H. Nordsieck, 2001
Cylindrophaedusa O. Boettger, 1877 - with three subgenera: Cylindrophaedusa and Montiphaedusa H. Nordsieck, 2002
Dautzenbergiella Lindholm, 1924 - with two subgenera: Dautzenbergiella and Mansuyiella H. Nordsieck, 2003
Diceratoptyx Pilsbry, 1905
真管螺属 Euphaedusa O. Boettger, 1877[22]： - with five subgenera: Dentiphaedusa H. Nordsieck, 2003; Euphaedusa; Papilliphaedusa H. Nordsieck, 2003; Tauphaedusa H. Nordsieck, 2003 and Telophaedusa H. Nordsieck, 2003：又名小煙管蝸牛屬。
Fuchsiana Gredler, 1887
拟管螺属 Hemiphaedusa O. Boettger, 1877：with 10 subgenera: Dendrophaedusa H. Nordsieck, 2002; Hemiphaedusa; Hemiphaedusoides H. Nordsieck, 2001; Hemizaptyx Pilsbry, 1905; Labyrinthiphaedusa H. Nordsieck, 2001; Margaritiphaedusa H. Nordsieck, 2001; Notoptychia Ehrmann, 1927; Pinguiphaedusa Azuma, 1982; Placeophaedusa Minato, 1994 and Selenophaedusa Lindholm, 1924
Heterozaptyx Pilsbry, 1906
Juttingia Loosjes, 1965 - with two subgenera: Juttingia and Pseudohemiphaedusa H. Nordsieck, 2002
Lindholmiella Ehrmann, 1927
Liparophaedusa Lindholm, 1924
Loosjesia H. Nordsieck, 2002
Luchuphaedusa Pilsbry, 1901 - with two subgenera: Luchuphaedusa and Nesiophaedusa Pilsbry, 1905
Macrophaedusa Moellendorff, 1883
Macrophaedusella H. Nordsieck, 2001
Megalophaedusa O. Boettger, 1877 - with four subgenera: Megalophaedusa; Mesophaedusa Ehrmann, 1929; Mesozaptyx Kuroda, 1963 and Mundiphaedusa Minato, 1979. (Neophaedusa is a synonym of Megalophaedusa).[23]
Metazaptyx Pilsbry, 1905
Miraphaedusa H. Nordsieck, 2005
Oligozaptx Pilsbry, 1905
Paraphedusa O. Boettger, 1877
Parazaptyx Pilsbry, 1905
Pliciphaedusa H. Nordsieck, 1998
Pulchraptyx Minato, 1981
Reinia Kobelt, 1876 - with three subgenera: Parareinia H. Nordsieck, 1998; Pictophaedusa Azuma, 1982 and Reinia
Renschiphaedusa Loosjes & Loosjes-van Bemmel, 1973
Selenoptyx Pilsbry, 1908
Serriphaedusa H. Nordsieck, 2001
Sinigena Lindholm, 1925
Stereozaptyx Pilsbry, 1905
Streptodera Lindholm, 1925
Synprosphyma Germain, 1919 - with two subgenera; Excussispira Lindholm, 1925 and Synprosphyma
Thaumatoptyx Pilsbry, 1908
Tyrannophaedusa Pilsbry, 1901 - with two subgenera: Aulacophaedusa Azuma, 1982 and Tyrannophaedusa
Tyrannozaptyx Käufel, 1930
Zaptychopsis Ehrmann, 1927
Serrulininae
Caspiophaedusa Lindholm, 1924
Dobatia H. Nordsieck, 1973
Graecophaedusa Rähle, 1982
Laeviphaedusa Likharev & Stelkov, 1965
Microphaedusa H. Nordsieck, 1978
Nothoserrulina Németh & Szekeres, 1995
Pamphylica Németh & Szekeres, 1995
Pravispira Lindholm, 1924
Pontophaedusa Lindholm, 1924
Pontophaedusella H. Nordsieck, 1994
Sciocochlea C. Boettger, 1935
Serrulina Mousson, 1873
Serrulinella H. Nordsieck, 1984
Truncatophaedusa Majoros, Németh & Szili-Kovács 1994
Tsoukatosia Gittenberger, 2000 - with two subgenera: Agiosspeleikos A. & P. Reischütz, 2003 and Tsoukatosia
保育
儘管非海洋軟體動物當中本科物種看似易危[24]，但在《國際自然保護聯盟瀕危物種紅色名錄》只有9個本科的物種被列入名錄內[25]。
參考文獻
^ J. E. Gray. April 14, 1855. Catalogue of Pulmonata or air-breathing Mollusca in the collection of the British Museum, Part I: 156.
^ 皱纹岛管螺. 科学数据库-中国动物主题数据库-中国动物物种编目数据库. [2018-06-11] （中文（简体））.
^ 3.03.13.23.3 宋大祥; 吳岷 (编). 英漢動物學詞典. 科學出版社. 2008. ISBN 978-7-03-021302-0 （中文（简体））.
^ Mesurethra. Integrated Taxonomic Information System. [2018-05-27] （英语）.
^ 5.05.15.25.35.45.55.6 蝸牛不思議
^ 陳德牛; 張國慶 (编). 德氏拟管螺 Hemiphaedusa draesekei Blume. 中国动物志. 无脊椎动物 第19卷 软体动物门 腹足纲 柄眼目 烟管螺科 第一版. 科学出版社. [2018-07-16]. ISBN 9787030067975 （中文（简体））.
^
The biology of terrestrial molluscs於Google Books。
^ 8.08.18.28.38.48.5 Frýda, J.; Hausdorf, B.; Ponder, W.; Valdés, Á.; Warén, A. Bouchet P.; Rocroi J.-P., 编. Classification and nomenclator of gastropod families. Malacologia: International Journal of Malacology (Hackenheim, Germany / Ann Arbor, USA: ConchBooks). 2005, 47 (1-2): 397 pp. ISBN 3-925919-72-4. ISSN 0076-2997.
^ Jörger, Katharina M; Stöger, Isabella; Kano, Yasunori; Fukuda, Hiroshi; Knebelsberger, Thomas; Schrödl, Michael. On the origin of Acochlidia and other enigmatic euthyneuran gastropods, with implications for the systematics of Heterobranchia. BMC Evolutionary Biology. 2010, 10: 323. PMC 3087543. PMID 20973994. doi:10.1186/1471-2148-10-323 （英语）.
^ Bouchet, P.; Rocroi, J.P.; Hausdorf, B.; Kaim , A.; Kano , Y.; Nützel , A.; Parkhaev, P.; Schrödl, M.; Strong , E.E. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia. 2017, 61 (1-2): 1–526. doi:10.4002/040.061.0201 （英语）.
^ Nordsieck H. (1981). Archiv für Molluskenkunde 111(1-3): 101.
^ Nordsieck H. (1978). Archiv für Molluskenkunde 109(1-3): 104.
^ Nordsieck H. (1985). Heldia 1(3): 83.
^ 14.014.114.214.314.414.514.614.714.8 "Clausiliini". Fauna Europaea, last update 27 January 2011, accessed 27 April 2011.
^ "Graciliaria". Fauna Europaea, last update 27 January 2011, accessed 27 April 2011.
^ Uit de Weerd D. R. (2004). "Molecular phylogenetic history of eastern Mediterranean Alopiinae, a group of morphologically indeterminate land snails". Doctoral thesis, Leiden University, ISBN 90-6464-874-3.
^ Gittenberger E. & Uit de Weerd D. R. (2009). "Summarizing data on the Inchoatia taxa, including Inchoatia megdova bruggeni subspec. nov. (Gastropoda, Pulmonata, Clausiliidae)" Zoologische Mededelingen 83 http://www.zoologischemededelingen.nl/83/nr03/a08
^ Family: CLAUSILIIDAE. WMSD - Worldwide mollusc species DB. [2018-04-02] （英语）.
^ 引用错误：没有为名为shelldoc/中国动物志W7的参考文献提供内容
^ 管螺亚科 Phaedusinae. 《中国动物志 无脊椎动物》 第七卷. [2018-04-02] （中文（简体））. [永久失效連結]
^ 21.021.1 Maassen W. J. M. & Gittenberger E. (2007). "Three new clausiliid land snails from Tonkin, northern Vietnam (Gastropoda: Pulmonata: Clausiliidae)". Zoologische Mededelingen 81(1): http://www.zoologischemededelingen.nl/81/nr01/a10
^ 真管螺属 Euphaedusa. 《中国动物志 无脊椎动物》 第七卷. [2018-04-02] （中文（简体））. [永久失效連結]
^ cf. Nordsieck H. (2006). "Species list of recent Clausiliidae".
^ Lydeard, C.; Cowie , R.; Ponder , W.F.; 等. The global decline of nonmarine mollusks (PDF). BioScience. 2004-04, 54: 321–330 [2009-10-20]. doi:10.1641/0006-3568(2004)054[0321:TGDONM]2.0.CO;2. （原始内容 (PDF)存档于2007-03-31） （英语）.|author=和|last1=只需其一 (帮助);|archiveurl=和|archive-url=只需其一 (帮助);|accessdate=和|access-date=只需其一 (帮助);|archivedate=和|archive-date=只需其一 (帮助)
^ IUCN 2009. IUCN Red List of Threatened Species. Version 2009.2. <www.iucnredlist.org>. Downloaded on 5 March 2010.
延伸閱讀
Maltz, T. K.; Sulikowska-Drozd, A. Life Cycles of Clausiliids of Poland — Knowns and Unknowns. Annales Zoologici. 2008, 58 (4): 857–880. doi:10.3161/000345408X396783 （英语）.- Nordsieck H. (2007). Worldwide Door Snails. ConchBooks, 213 pp. ISBN 978-3-939767-07-7.
- Uit de Weerd D. R. (2004). "Molecular phylogenetic history of eastern Mediterranean Alopiinae, a group of morphologically indeterminate land snails". Doctoral thesis, Department of Biology, Faculty of Science, Leiden University. HTM, PDF.
- Uit de Weerd D. R., Piel W. H. & Gittenberger E. (2004). "Widespread polyphyly among Alopiinae snail genera: when phylogeny mirrors biogeography more closely than morphology". Molecular Phylogenetics and Evolution 33(3): 533-548. doi:10.1016/j.ympev.2004.07.010
外部連結
 | 维基共享资源中相关的多媒体资源：烟管蜗牛科 |
- Nordsieck H. (2005–2007). www.clausilia.de at Internet Archive.
- Nordsieck H.: http://www.hnords.de: Internet journal for the malacology of landsnails (with special regard to Clausiliidae).
- BioLib info
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